Margaery Tyrell is the only daughter of Alerie Tyrell (neé Hightower) and Mace Tyrell the Lord of House Tyrell, one of the Great Houses of the Seven Kingdoms. She is the sister of Ser Loras Tyrell, Garlan Tyrell and Willas Tyrell. She is the granddaughter of Lady Olenna Redwyne. The Tyrells' seat of power is Highgarden, capital of the Reach.
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